Animations flickering during playtest but not in animation editor

I’ve animated a custom bird model, but for some reason the animation seems to flicker and snap randomly during testing, and not when it is is previewed in the animation editor (videos below).
This issue happens both when the animation is played locally and when it is replicated from the server.

In editor / in game:


Extra info about animation / rig:

  • Animation priority: Movement
  • Rig uses an AnimationController (is not a humanoid).
  • Rig is structured as shown below (rigs in both clips are identical):

    (Excuse the use of unions; they are placeholders and will be replaced later.)

Does anyone know what could be causing this issue / how to fix it?

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